What the habitat amount hypothesis does and does not predict: A reply to Saura

Saura (2020) used the habitat amount hypothesis (HAH) to predict negative effects of fragmentation per se on mean species density site over a region. This prediction is valid but incomplete; HAH can also positive stated, "the compatible with steeper slope species–area relationship for fragmented than continuous habitat, and higher richness single large patch several small patches same total area (SLOSS)." Importantly, does not species-area (SAR) slopes or SLOSS. These require information about how composition changes space, while only predicts site. The therefore equally shallower SAR either outcome